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Contents
Family Delphacidae Leach, 1815
Subfamily Delphacinae Leach, 1815
Tribe Delphacini Leach, 1815
Genus Delphacodes Fieber, 1866: 524.
Type species (In original combination): Delphax mulsanti Fieber, 1866: 526 (designated by Kirkaldy 1904: 177).
Distribution
Delphacodes in the strict sense is a western Palearctic genus (with 10 species); however, currently there are species placed in this genus from all over the world.
Distribution of Delphacodes from FLOW (as of 20 Feb. 2020)
Recognized species
By my count (below), there are 157 species currently placed in this genus. However, following Asche & Remane (1983), there are only 10 species legitimately placed in this genus.
There has long been confusion regarding the definition and composition of Delphacodes. The genus is discussed (among others) by Muir & Giffard (1924), Haupt (1935), China (1954), Dlabola (1957, 1961), Linnavuori (1957), Nast (1958), Wagner (1963) and Le Quesne (1964). The modern definition of Delphacodes follows Asche & Remane (1983).
Species list: Delphacodes sensu stricto– Distribution information for Old World species may be incomplete.
Distribution information may be incomplete.
1 Delphacodes audrasi Ribaut, 1954 – France
2 Delphacodes capnodes (Scott, 1870: 69) – Austria, Bulgaria, Czech Republic, Estonia, France, Germany, Great Britain, Greece, Hungary, Netherlands, Poland, Serbia, Sweden, Yugoslavia
3 Delphacodes fascia (Lindberg, 1960) – Portugal
= Calligypona fascia Lindberg, 1960 previous combination of Delphacodes fascia (Lindberg, 1960) by Asche & Remane (1983: 31)
4 Delphacodes framarib Asche and Remane 1983 – Spain
5 Delphacodes linnavuori (Le Quense, 1960) – France
= Delphacodes mulsanti Linnavuori 1957 and Dlabola (1957) nec Fieber 1866
= Megamelodes linnavuorii Le Quesne 1960: 186c
= Delphacodes linnavuorii (Le Quesne); comb. by Dlabola 1981: 156, Asche & Remane, 1983 but see Nast 1972
6 Delphacodes mulsanti (Fieber, 1866: 526) – France
7 Delphacodes nastasiAsche and Remane 1983 – Greece
8 Delphacodes ornatipennis (Haupt, 1927) – Afghanistan, Iran, Israel, Jordan, Palestine
= Megamelus ornatipennis Haupt, 1927 (original combination) to Delphacodes ornatipennis (Haupt, 1927) by Asche & Remane (1983)
9 Delphacodes schinias Asche and Remane 1983 – Greece
10 Delphacodes venosus (Germar, 1830) – Albania, Austria, Belgium, Bulgaria, Czech Republic, Denmark, Estonia, Finland, France, Germany, Great Britain, Greece, Hungary, Italy, Latvia, Moldova, Netherlands, Norway, Poland, Russia (North European Russia), Sweden, Switzerland, Ukraine, Yugoslavia
= Delphax venosus Germar, 1830. original combination
= Delphax rhyparia Flor, 1861; syn by Fieber 1872: 6.
= Delphax piceola Kirschbaum, 1868; syn. by Wagner, 1939: 119.
= Liburnia melanopachys Scott, 1870; syn by Oshanin 1907: 322, noted Edwards 1924: 57.
= Liburnia curtula Sahlberg, 1871; syn. by Puton 1875: 120.
= Liburnia thoulessi Edwards, 1894; syn by Oshanin 1907: 322 (notes also China 1938: 243).
= Delphax nitidula Kirschbaum, 1868; syn by Fieber 1872: 6.
Species list: Delphacodes sensu lato
These species are currently placed in Delphacodes, but their actual status is unclear. It is entirely possible that some of these names represent missed synonyms on my part.
Palearctic species of uncertain status – Fauna Europaea treats many of these species under their original combination (as noted).
1 Delphacodes agropyri Ishihara, 1949: 55- Japan (Honshu)
2 Delphacodes axillaris (Sahlberg, 1876: 138) – Finland (Fauna Europaea places this species in Liburnia).
3 Delphacodes burjata (Kusnezov, 1929) – Turkestan
4 Delphacodes conspicua (Horvath, 1904) – Turkestan, Kazakhstan
5 Delphacodes esakii Matsumura & Ishihara, 1945: 63- Japan, Taiwan
6 Delphacodes flava Metcalf, 1943 (new name for Delphax flavescens Ivanoff 1885, not Fabricius 1794: 7) – Russia, Ukraine (see Logvinenko 1975)
7 Delphacodes fukuokae Ishihara, 1949: 57- Japan (Kyushu)
8 Delphacodes fuscifrons (Fieber, 1879: 92) – Belgium, France (Fauna Europaea places this species in Liburnia)
9 Delphacodes gilveola (Kirschbaum, 1868) – Austria, Germany, Turkestan (Fauna Europaea places this species in Delphax)
10 Delphacodes gracilis (Matsumura, 1915) – Korea
11 Delphacodes hemiptera (Germar, 1818) – Europe (reported in error: Alaska; Fauna Europaea places this species in Delphax)
12 Delphacodes latifrons (Puton 1875) (= Liburnia latifrons Fieber, 1872 nom. nud.) – France, Germany (Fauna Europaea places this species in Liburnia)
13 Delphacodes leptypha (Amyot, 1847) – France
14 Delphacodes macroptera(Costa, 1834) – Italy (Fauna Europaea places this species in Delphax)
15 Delphacodes melanocephala (Fieber, 1879: 101) (incertae sedis)- Austria, Germany, Russia, Ukraine
16 Delphacodes modesta (Fieber, 1866: 526) – France (Fauna Europaea places this species in Delphax)
17 Delphacodes nigerrima Ishihara, 1949 – Japan (Honshu)
18 Delphacodes radiata (Costa, 1834) – Italy (Fauna Europaea places this species in Delphax)
Delphacodes sahlbergiana Ishihara, 1949: 66 (replacement name for Calligypona albicollis Sahlberg, 1871: 409); jr. syn. of Calligypona albicollis by Fujinama (2013: 246) [Metcalf 1943]
Delphacodes shirozui Ishihara, 1949: 53-54 (see Metadelphax propinqua); syn. with D. propinqua (Fieber) by Fennah 1956b: 122.
19 Delphacodes stigmaticalis (Curtis, 1837) – England, Ireland, Great Britain (Fauna Europaea places this species in Delphax)
20 Delphacodes tapina (Fieber, 1866: 528) – Denmark, France, Italy (Sardinia) (Fauna Europaea places this species in Delphax)
21 Delphacodes uncinata (Fieber, 1866: 524) – Great Britain, Greece, Hungary, Portugal (Fauna Europaea places this species in Delphax)
22 Delphacodes walkeri Metcalf, 1943: 522 – Egypt
= Delphax dorsalis Walker 1870 (nec Fitch 1851)
Palearctic species ‘recently’ in Delphacodes now placed in other genera
Delphacodes albifrons(Fieber, 1879) now placed in Mirabella Emeljanov 1982
Delphacodes akashiensis (Matsumura, 1900) is a jr syn of Laodelphax striatellus (Fallén, 1826: 75), synonymized by Matsumura & Ishihara 1945: 61.
Delphacodes albocarinata (Stal, 1858) (and subspecies) — moved to Tyrphodelphax albocarinatus (Stål, 1858) by Vilbaste 1968, then to Nothodelphax albocarinata (Stål, 1858) by Anufriev & Averkin (1982): 137 (subspecies Delphacodes albocarinata intermedia (Sahlberg, 1871) & Delphacodes albocarinata palliata (Sahlberg, 1871) should follow to Nothodelphax); subspecies palliata was treated as a jr syn of albocarinata by Wagner 1963; and intermedia as jr syn of albocarinata by Nast (1972).
Delphacodes alboguttata (Melichar, 1903) is now Nanotoya alboguttata (Melichar, 1903) as the type species of Nanotoya Fennah 1975.
Delphacodes alpina (Sahlberg, 1871: 462) and subspecies [Delphacodes alpina tota (Sahlberg, 1871), Delphacodes alpina nigrovariegata (Sahlberg, 1871), Delphacodes alpina grisescens (Sahlberg, 1871), Delphacodes alpina griseoflavescens (Sahlberg, 1871), Delphacodes alpina dorsoflava (Sahlberg, 1871)] belong in Javesellabased on Ossiannilsson (1978: 200); all treated as junior syn of Javesella forcipata by Nast 1972: 67.
Delphacodes anemonias (Kirkaldy, 1907) is now the type species of Terthron Fennah 1965: 55-56 (the Australia paper)
Delphacodes biarmica (Sahlberg, 1871) is a jr syn of Ribautodelphax collinus (Boheman, 1847) made by (or before) Wagner 1963.
Delphacodes boldi (Scott, 1870) was placed in Gravesteiniella Wagner, 1963 by Wagner 1963.
Delphacodes bohemani (Stål, 1858) (nec Stal 1854) was replaced by Delphacodes stali Metcalf, 1943.
Delphacodes brevipennis (Boheman 1847) (and subspecies D. b. sordidella (Sahlberg, 1871) were moved to Muellerianella brevipennis (Boheman, 1847) by Wagner 1963.
Delphacodes capellana (Jacobi, 1917) was moved to Syndelphaxby Fennah 1964: 143 (the Madagascar paper).
Delphacodes cataniae (Matsumura, 1910) is a jr syn of Metadelphax propinqua (Fieber) according to Nast (1975: 4).
Delphacodes clypealis (Sahlberg, 1871) is now in Paraliburnia Jensen-Haarup, 1917; This species was included by Wagner (1963) in this genus, but the transfer might have been done earlier (e.g., Le Quesne 1960a).
Delphacodes conwentzi Matsumura, 1906 is a junior synonym of Kosswigianella denticauda (Boheman, 1847) according to Wagner (1963).
Delphacodes dalei(Scott, 1870) was moved to the new genus Lamprodelphax by Fennah, 1982, subsequently, the genus was made a syn. of Scottianella Anufriev, 1980.
Delphacodes difficilis (Edwards, 1888) is a jr synonym of Javesella (Javesella) dubia (Kirschbaum, 1868) according to Wagner 1939: 125; Le Quesne 1960: 43. [Metcalf]
Delphacodes discreta (Edwards, 1888); junior synonym of Javesella (Javesella) obscurella (Boheman, 1847) by Wagner 1939: 125 (as Liburnia obscurella); Le Quesne 1960: 43.
Delphacodes distincta (Flor, 1861) moved to Nothodelphax distinctus (Flor) by Le Quesne 1964: 57.
Delphacodes distingueda (Kirschbaum, 1868), syn. with Ribautodelphax albostriata (Fieber, 1866) apparently by Wagner 1939.
Delphacodes dolosa Muir, 1926; to Syndelphax dolosa (Muir, 1926); comb. by Gonzon and Bartlett, 2008: 252.
Delphacodes exigua (Boheman, 1847), to Kosswigianella exigua (Boheman) by Wagner 1963 (type of genus) to Muirodelphax exiguus (Boheman). comb. by Le Quesne, 1964: 57 (Koswigianella subsumed under Muirodelphax); to Kosswigianella exigua (Boheman) by (e.g.) Vilbaste 1971: 138 or before.
Delphacodes fairmairei (Perris, 1857); made type species of Muellerianella by Wagner 1993: 175.
Delphacodes faimairei signicolli (Rey, 1894) to Muellerianella by Wagner (1963) with nomative subspecies (apparently subsumed under Muellerianella fairmairei (Perris) by Wagner (1963: 175)).
Delphacodes fallax Muir, 1926; to Syndelphax fallax (Muir, 1926); comb. by Gonzon and Bartlett, 2008: 252.
Delphacodes fieberi (Scott, 1870); to Megamelodes by Le Quesne 1960b: 3; syn. with Megamelodes quadrimaculatus (Signoret 1865) syn. by Wagner 1963: 175.
Delphacodes flaveola (Flor, 1861), to Xanthodelphax (as type species) by Wagner, 1963: 176).
= Delphacodes flaveola dorsoinfuscata Metcalf, 1943; jr syn of Xanthodelphax flaveolus (Flor) by Wagner 1963: 176.
Delphacodes flavipennis (Sahlberg, 1871) (original combination Liburnia flavipennis Sahlberg, 1871); syn of Javesella pellucida by Wagner 1963: 176.
Delphacodes forcipata (Boheman, 1847), to Weidnerianella Wagner 1963: 176, then to Javesella by Le Quesne 1964: 57 & Wagner 1966: 91.
Delphacodes frontalis (Kirschbaum, 1868); jr synonym of Javesella pellucida by Wagner 1963: 176.
Delphacodes fumipennis (Fieber, 1866), type species of Flastena Nast 1975 (as Flastena fumipennis (Fieber, 1866)) by original designation (jr. syn of Javesella discolor (as Liburnia) in Wagner 1939: 125 [error]).
Delphacodes fuscovaria (Distant, 1917); type species of Opiconsiva Distant, 1917: 301 (as Opiconsiva fuscovaria Distant, 1917: 301 ) by original designation.
Delphacodes giffuensis (Matsumura, 1900); given as jr syn of Laodelphax striatellus in Takara (1959: 5), syn. in Matsumura & Ishihara (1945: 61).
Delphacodes guaramanensis (Scott, 1873); jr. syn. of Florodelphax leptosoma by Webb & Wilson (1986: 285); but Nast 1986: 229 made this species a jr syn of Scottianella dalei.
Delphacodes haglundi (Sahlberg, 1871); moved to Megadelphax by Ossiannilsson 1978: 115; see also Emeljanov 1982: 85 (“Calligyphona” haglundi (Sahlberg) according to Dlabola 1955: 122; Tyrphodelphax haglundi (Sahlberg, 1871); according to Nast 1972: 60; sr syn of Megadelphax paulus Emeljanov, 1972).
Delphacodes herrichii (Kirschbaum, 1868); jr syn. of Javesella pellucida in Wagner 1963: 176 (but jr. syn of Javesella dubia in Wagner 1939: 125).
Delphax hyas Kirkaldy 1907a (= Delphacodes hyas (Kirkaldy), comb. by Muir 1917: 333), jr synonym of Syndelphax disonymos (Kirkaldy) by Bellis & Donaldson 2015: 10.
Delphacodes juncea (Haupt, 1935); comb. by Metcalf 1943: 456; syn. with Javesella salina (Haupt) by Wagner 1963: 176.</>
Delphacodes leprotosoma (Flor, 1861) (possibly a misspelling for Delphax leptosoma Flor 1861)
Delphacodes leptosoma (Flor, 1861), to Florodelphax Vilbaste 1968b in Vilbaste (1968b – the key to nymphs) (= Liburnia guaramensis Scott, 1873; syn. by Webb & Wilson 1986; Delphax heydenii Kirschbaum, 1868, syn by Fieber, 1872: 450, 465; Delphax albofimbriata Fieber, 1866, syn by Fieber, 1872: 450, 465).
Delphacodes lethierryi (Scott, 1873: 105) (Corsica); syn. with Flastena fumipennis (Fieber, 1866) by Webb & Wilson 1986.
Delphacodes limitata (Fieber, 1866); jr syn of Kosswigianella exigua in Wagner 1963: 175.
Delphacodes lucticolor (Sahlberg, 1868), syn of Florodelphax paryphasma (Flor, 1861) by Ossiannilsson 1978: 156.
Delphacodes lugubrina (Boheman, 1847), type species of Struebingianella Wagner 1963 by original designation.
Delphacodes maikoensis (Matsumura, 1900: 266), syn with Laodelphax striatellus (as Delphacodes striatella) by Matsumura and Ishihara 1945: 61.
Delphacodes marshalli (Scott, 1873), syn. with Metadelphax propinqua (as Toya) by Webb & Wilson 1986: 286.
Delphacodes minoensis (Matsumura, 1900) (Japan), syn with Laodelphax striatellus (as Delphacodes striatella) by Matsumura and Ishihara 1945: 61.
Delphacodes mocsaryi (Horváth, 1897) (Hungary); junior syn of Kosswigianella exigua by Asche 1982 (the Geza Horvath paper).
Delphacodes nigeriensis (Muir, 1920); to Sogatella nigeriensis (Muir, 1920) by Asche and Wilson 1990: 13
Delphacodes nigricans (Matsumura, 1910); syn. with Flastena fumipennis (Fieber, 1866) by Nast 1975.
Delphacodes nigrifrons (Matsumura, 1910), to Matutinus putoni (Costa, 1888) by Asche & Remane 1982.
Delphacodes nipponica (Matsumura, 1900), syn with Laodelphax striatellus (as Delphacodes striatella) by Matsumura and Ishihara 1945: 61.
Delphacodes nitidipennis (Kirschbaum, 1868); a jr syn of Javesella dubia (as Liburnia dubia) by Wagner 1939: 125.
Delphacodes niveimarginata (Scott, 1870), syn. with Florodelphax paryphasma (as Delphacodes) in Vilbaste 1960, 1968; also Webb & Wilson 1986: 286.
Delphacodes obscurella (Boheman, 1847), to Weidnerianella obscurella (Boheman, 1847) by Wagner 1963: 176, then to Javesella obscurella (Boheman, 1847)by Le Quesne 1964: 57 & Wagner 1966: 91.
Delphacodes obscurinervis (Kirschbaum, 1868), syn. with Muirodelphax aubei by Wagner, 1963: 175
Delphacodes pallecips (Horváth, 1897); missp. for palliceps (Horváth, 1897 (in FLOW)).
Delphacodes palliceps (Horváth, 1897: 623); type species of Horvathianella Anufriev, 1980: 214 by original designation.
Delphacodes pallens (Stål, 1854); transferred to Ribautodelphax Wagner 1963 by Wagner 1963.
Delphacodes paludicola (Muir, 1926) (in FLOW) should be Delphacodes paludicola Muir, 1926 (here in Metcalf 1943)
Delphacodes paludosa (Flor, 1861); type species by monotypy and original designation of ParadelphacodesWagner 1963.
Delphacodes parvistylus Muir, 1929 (South Africa)
Delphacodes pellucida fuscicollis (Rey, 1891: 244); listed as synonym of Javesella pellucida (Fab.) (as Delphacodes) in Esaki & Ishihara 1945: 31. (Metcalf 1943)
Delphacodes pellucida obscura (Fieber, 1872); listed as synonym of Javesella pellucida (Fab.) (as Delphacodes) in Esaki & Ishihara 1945: 31. (Metcalf 1943)
Delphacodes pellucida varicollis (Rey, 1894: 14); listed as synonym of Javesella pellucida (Fab.) (as Delphacodes) in Esaki & Ishihara 1945: 31. (Metcalf 1943)
Delphacodes protrusa (Fieber, 1866), syn with Muirodelphax aubei by Vilbaste 1971: 146.
Delphacodes reyi (Fieber, 1866), to Calligypona by Wagner 1963: 175; Le Quesne 1964: 56-57 (= Delphax longipes Kirschbaum, 1868, = Calligypona albicollis Sahlberg, 1871, =Delphacodes sahlbergiana Ishihara, 1949: 66 (replacement name for Calligypona albicollis Sahlberg, 1871: 409); jr. syn. of Calligypona albicollis by Fujinama (2013: 246) [Metcalf 1943]
Delphacodes scutellata (Scott, 1873), syn. with Kosswigianella exigua by Le Quesne 1960: 14, 19.
Delphacodes signoreti (Scott, 1870), syn. with Paraliburnia adela (Flor) by Le Quesne 1960: 51 (as Delphacodes), see also Webb & Wilson 1986: 267.
Delphacodes similis (Kirschbaum, 1868); jr syn of Javesella discolor in Wagner 1939: 125; Nast 1972: 66.
Delphacodes simulans (Walker, 1851) (Galapagos), moved into Nesosydne by Fennah 1967: 67 (the galapagos paper).
Delphacodes sordida (Kirschbaum, 1868), jr syn of Paraliburnia adela (Flor, 1861) in Wagner 1963: 175.
Delphacodes spinigera Fennah, 1945, to Pygospina spinigera (Fennah, 1945); comb. by Caldwell and Martorell, 1951: 169.
Delphacodes stali Metcalf, 1943: 510, nom. nov. for Delphax bohemani Stal 1858 [nec Delphax bohemani Stal 1854 (Cixiidae).]
Delphacodes straminea (Stål, 1858), to Xanthodelphax by Wagner 1963; (= Delphax fulveola (Kirschbaum 1868 according to Wagner 1939, 1963 [syn. of flaveola Flor elsewhere]; = Liburnia v-flava Scott 1881 by Edwards, 1886: 86)
Delphacodes striatella fimbriata (Rey, 1894: 14), syn. with Laodelphax striatella (as Callidelphax Wagner 1963 – Laodelphax striatella is type species of Callidelphax Wagner 1963 by monotypy and original designation).
Delphacodes strigosa(Matsumura, 1910), syn of Toya suezensis (Matsumura, 1910) by Nast 1975: 5 (Toya suezensis is listed as a jr syn of Flastena fumipennis (Fieber, 1866) in Fauna Europaea, but I have not found where this syn. has occurred, yet.).
Delphacodes thomasseti(Muir, 1925) To Toya thomasseti (Muir, 1925) comb. by Fennah (1964: 141) ; (= Opiconsiva modesta Distant 1917, Delphacodes distanti Metcalf, 1943, noted in Fennah 1964: 141, the Madagascar paper, syns. not noted as new).
Delphacodes thoracica (Distant, 1916); to Harmalia thoracica (Distant), comb. by Fennah 1969: 39 (Type species of genus Harmalia))
Delphacodes tschikoica (Kusnezov, 1929) – Turkistan ((= Liburnia tschikoica Kusnezov, 1929, original combination, to Delphacodes by Metcalf 1943: 520); syn of Gravesteiniella boldi (Scott, 1870) by Vilbaste 1965: 12; see also Vilbaste 1968: 30, where this appears to be a subspecies of Gravesteiniella boldi (Scott, 1870).
Delphacodes unicolor (Walker, 1851), replaced by Delphacodes uniformis Metacalf, 1943 (insertae sedis, adescribed from a nymph)
Delphacodes vegetatus (Melichar, 1912: 133) – Belgian Congo, Ziare (= Peregrinus vegetatus Melichar, 1912; to Delphacodes by Muir 1926: 28). Now in Thriambus by Fennah 1964: 144 [the Madagascar paper])
Australian species of uncertain status
Delphacodes hyas (Kirkaldy, 1907) – Queensland – Now junior syn of Syndelphax disonymos (Kirkaldy) by Bellis and Donaldson 2015: 10.
African species of uncertain status (including nearby islands)
Delphacodes actaeon Fennah, 1958 – (Mauritania) to Toya actaeon (Fennah, 1958) by Linnavuori (1973: 107)
Delphacodes aglauros Fennah, 1958 (Senegal) see Falcotoya
1 Delphacodes anthracina (Horvath, 1909: 301) – Canary Islands, Maderia Archipelago (Fauna Europaea places this species in Liburnia)
2 Delphacodes catilina Fennah, 1958 – SW Africa (called “Toya catilina” by Harder and Bakker 1973, Abo and Sy 1997)
3 Delphacodes distanti Metcalf, 1943 – Seychelles
4 Delphacodes graminis Lindberg, 1958 – Cape Verde Islands
5 Delphacodes parvistylus Muir, 1929 – South Africa
6 Delphacodes sabrina Fennah, 1958 – South Africa
7 Delphacodes semiobscura Metcalf, 1943 (new name Liburnia seminigra Melichar 1905, not Delphax seminigra Stal 1854) – Tanganyika
Indo-Malayan species of uncertain status
Delphacodes elegantissima lshihara, 1952: 45; jr. syn of Sogatella kolophon
1 Delphacodes kotonis (Matsumura, 1940: 49) – Taiwan
= Liburnia kotonis Matsumura, 1940; to Delphacodes by Ishihara 1949: 58.
2 Delphacodes flavida(Melichar, 1903) – Sri Lanka
Delphacodes furcifera (Horváth 1899; 372) – see Sogatella
3 Delphacodes kahavalu Kirkaldy 1906: 156 (new name for Delphax venosus Motschulsky, 1863 nec Gemar) – Sri Lanka
= Liburnia ceylanica Jensen-Haarup, 1920, new name for Delphax venosus De Motschulsky, 1863; synonym of Delphacodes kahavalu (Kirkaldy, 1906) in Metcalf 1943: 456.
4 Delphacodes marginalis (Motschulsky, 1863) – Sri Lanka
5 Delphacodes pallidula (Melichar, 1903) – Sri Lanka
= Liburnia pallidula Melichar 1903: 103 (original combination)), to Delphacodes in Metcalf 1943: 480.
= Delphacodes taprobanensis Kirkaldy, 1906 (new name for Liburnia pallidula Melichar), syn. in Metcalf 1943: 480.
6 Delphacodes unistrigosa (Motschulsky, 1863) – Sri Lanka
Pacific Island species of uncertain status
1 Delphacodes amblystylis Fennah, 1956 – Mariana Island, Guam
2 Delphacodes celaeno Fennah, 1956 – Caroline Island
3 Delphacodes choroebus Fennah, 1958 – Mangareva Island
4 Delphacodes muirella Metcalf, 1943 (new name for Delphacodes pacifica Muir 1927, not Megamelus pacificus Crawford 1914: 626)- Tahiti
5Delphacodes muirianus Izzard, 1936 – Christmas Island
6 Delphacodes thersander Fennah, 1956 – Caroline Island
1 Delphacodes ixionFennah, 1958 – Cook Island
The range information is intended to be a full compilation of published range information, although I may have missed some entries. US State abbreviations follow the US Postal Service. Canadian Province abbreviations follow Maw et al. (2000).
1 Delphacodes aculeata Beamer, 1948 – USA: FL – Jr synonym of Chionomus puellus synonymy by Weglarz and Bartlett 2020: 31.
2 Delphacodes acuministyla Dozier, 1926 -USA : CT, IL, LA, MO, MS, VA
Delphacodes adunca Beamer, 1948b: 98, 105; to Flavoclypeus aduncus (Beamer 1948b) by Kennedy and Bartlett 2014: 24.
Delphacodes aetocephala Beamer, 1948b: 97; to Aethodelphax aetocephalus (Beamer, 1948b) Bartlett & Hamilton 2011: 54–55.
Delphacodes alata Beamer, 1948c: 118; to Aethodelphax alatus (Beamer, 1948c). by Bartlett & Hamilton 2011: 55–56.
Delphacodes albinotata Muir & Giffard, 1924: 36 (nec. Crawford 1914), replaced by Delphacodes arcuata Beamer, 1948
Delphacodes albinotata (Crawford, 1914: 619) (nec Muir & Giffard, 1924); jr syn. of Peregrinus maidis syn. by Beamer 1948b: 106.
Delphacodes albocarinata (Stål, 1858: 357); to Calligypona albocarinata (Stål, 1858) by Ossiannilsson 1946: 57; then to Muirodelphax albocarinata (Stål, 1858) by Wagner 1963: 175; then to Tyrphodelphax albocarinatus (Stål, 1858) by Vilbaste 1968: 70; 1971: 187; then Nothodelphax albocarinata (Stål, 1858) by Anufriev & Averkin 1982b: 137 (emendation to albocarinatus by Wilson 1992: 85–86.
Delphacodes albostriata (Fieber, 1866b: 525); to Ribautodelphax albostriatus (Fieber, 1866b) by Wagner 1963: 176.
Delphacodes alexanderi (Metcalf, 1923) – See Syndelphax
Delphacodes analis (Crawford, 1914: 620); to Calligypona analis (Crawford, 1914) by Kontkanen 1958: 143; then to Acanthodelphax analis (Crawford, 1914) by Wilson 1988: 338–339; then to Kosswigianella (Acanthodelphax) analis (Crawford, 1914) by Hamilton, 2002a: 20.
Delphacodes andromeda (Van Duzee, 1907: 46); to Flavoclypeus andromedus (Van Duzee, 1907) by Kennedy and Bartlett 2014: 24.
3 Delphacodes angulata Beamer, 1947 – USA: KS, MO, TX
4 Delphacodes anufrievi Wilson, 1992 – Canada: BC, SK, YK
5 Delphacodes apicata Beamer, 1948 – USA: AZ
6 Delphacodes arcuata Beamer, 1948 (jr. synonym of Chionomus tenae by Weglarz and Bartlett 2020: 36.)- USA: FL; Dominica, Guadeloupe, Mexico, Puerto Rico (inc. Vieques Is.)
= Delphacodes albinotata Muir & Giffard, 1924: 36 (nec Crawford, 1914) (D. arcuata is replacement name)
Delphacodes arcanastyla Beamer, 1948b: 101; to Javesella arcanastyla (Beamer, 1948b) by Wilson 1988: 341.
Delphacodes ardentis Beamer, 1948 – jr syn. Tagosodes wallacei
7 Delphacodes argentina De Remes, Lenikov & Teson, 1979 – Argentina
Delphacodes arvensis (Fitch, 1856: 46); to Muirodelphax arvensis (Fitch) by Hamilton & Kwon 2010: 423.
8 Delphacodes aterrima Muir, 1926 – Brazil (Pará), British Guiana (Surinam?), Colombia
Delphacodes atralabis Beamer, 1948b: 103; to Muirodelphax atralabis (Beamer, 1948b) by Hamilton & Kwon 2010: 422.
Delphacodes atrata Osborn, 1938: 344; to Javesella atrata (Osborn, 1938) by Wilson 1988: 341.
Delphacodes atridorsum Beamer, 1947: 63; to Caenodelphax atridorsum (Beamer, 1947) by Hamilton 2002a: 17; then to Flavoclypeus atridorsum (Beamer, 1947) by Kennedy and Bartlett 2014: 27.
9 Delphacodes atrior (Fowler, 1905: 134) – Mexico
Delphacodes axonopi Fennah, 1945a: 434 – jr. syn. of Toya nigra, see Toya
Delphacodes balboae Muir & Giffard, 1924: 36; to Chionomus balboae (Muir & Giffard, 1924) by Fennah 1971a: 324.
10 Delphacodes balli Muir and Giffard, 1924 – USA: CT; CAN: ON
11 Delphacodes banosensis Muir, 1926 – Ecuador (to Chionomus banosensis (Muir, 1926), combination by Weglarz and Bartlett 2020: 15)
Delphacodes basivitta (Van Duzee, 1909: 202); to Isodelphax basivitta (Van Duzee, 1909) by Fennah 1963a: 16
12 Delphacodes bellicosa Muir and Giffard, 1924 – USA: CA (to Chionomus bellicosus (Muir & Giffard, 1924), combination by Weglarz and Bartlett 2020: 16)
13 Delphacodes bergi (Scott, 1881: 156) – Argentina
Delphacodes bifida Beamer, 1948b: 99; to Elachodelphax (Aschedelphax) bifida (Beamer, 1948b) by Hamilton 2002a: 17.
Delphacodes bifurca Beamer, 1946 – see Yukonodelphax
14 Delphacodes bocana Beamer, 1946 – USA: TX
Delphacodes boxi Muir, 1926: 32; to Toya by Gonzon & Bartlett 208: 217.
15 Delphacodes caerulata Beamer, 1947 – USA: KS, MO
Delphacodes campestris (Van Duzee, 1897: 254); jr. syn. of Muirodelphax arvensis (Fitch, 1851) see Muirodelphax
Delphacodes canina Beamer, 1947: 60; to Aethodelphax caninus (Beamer, 1947) by Bartlett & Hamilton 2011: 56–57.
16 Delphacodes carinata (Glover, 1877) – USA (nomen dubium)
17 Delphacodes cayamensis (Crawford, 1914: 614) – Cuba
18 Delphacodes cerberus Fennah, 1957 – Juan Fernández Islands
19 Delphacodes chiloensis Muir, 1929 – Chiloe Is. (Chile)
Delphacodes coloradensis Beamer, 1952a: 114; to Aschedelphax coloradensis (Beamer, 1952a) by Wilson 1992: 89–90; then Elachodelphax (Aschedelphax) coloradensis (Beamer, 1952) Hamilton 2002a: 17.
Delphacodes compta Beamer, 1948c; to Akemetaopon comptum (Beamer, 1948c) by Bartlett et al. 2014: 109.
Delphacodes concava Beamer, 1948c: 113; to Aethodelphax concavus (Beamer, 1948c) by Bartlett & Hamilton 2011: 57–59.
Delphacodes consimilis (Van Duzee, 1897: 249); to Nothodelphax consimilis (Van Duzee, 1897) by Fennah 1965d: 254.
Delphacodes cornuta Beamer, 1948 – See Euides.
20 Delphacodes culta (Van Duzee, 1907: 47) – USA: FL: Bermuda, Cuba, Jamaica. (to Chionomus cultus (Van Duzee, 1907), incertae sedis combination by Weglarz and Bartlett 2020: 18.)
21 Delphacodes curvistyla Dozier, 1926 – USA: LA, MS
22 Delphacodes darwini Muir, 1929 – Chiloe Is. (Chile)
Delphacodes dentipennis Beamer, 1948b; syn. withIsodelphax nigridorsum (Crawford, 1914) by Bartlett et al. 2014: 118.
23 Delphacodes dentis Beamer, 1948 – USA: TX; CAN: NT (refuted) (synonymy with Chionomus dissipatus (Muir, 1926) by Weglarz and Bartlett 2020: 19).
Delphacodes detecta (Van Duzee, 1897: 248); to Spartidelphax detectus (Beamer, 1950) by Bartlett and Webb 2014: 25.
Delphacodes discolor (Boheman, 1847: 61); to Javesella discolor (Boheman, 1847) by Le Quesne 1964: 57.
Delphacodes dolera (Spooner, 1912: 240); to Javesella dolera (Spooner, 1912); comb. by Maw et. al. 2000: 84.
24 Delphacodes elongatus Teson and Remes Lenicov, 1983 – Argentina
Delphacodes emeljanovi Wilson, 1992; to Kosswigianella emeljanovi (Wilson, 1992) by Bartlett et al. 2014: 121.
Delphacodes excisa (Melichar, 1898: 67) – see Unkanodes
Delphacodes fallax Muir, 1926: 33 – See Syndelphax
Delphacodes foveata (Van Duzee, 1897: 257); to Nothodelphax foveata (Van Duzee, 1897) by Fennah 1963a: 15 (designated type species of Nothodelphax).
Delphacodes foveata subfoveata Muir & Giffard, 1924: 22; to Nothodelphax foveata subfoveata (Muir & Giffard, 1924) by implication Fennah 1963a: 15.
Delphacodes floridae Muir and Giffard, 1924 – See Syndelphax
Delphacodes floridae puertoriciensis Caldwell, 1951 – See Syndelphax
Delphacodes fulvidorsum Metcalf, 1923) – See Syndelphax
25 Delphacodes furcata (Provancher, 1872) (incertae sedis) – USA: NH, NY; CAN: ON, QC
Delphacodes gillettei (Van Duzee, 1897: 258); to Nothodelphax gillettei (Van Duzee, 1897) by Fennah 1965d: 254.
26 Delphacodes gluciophila Muir, 1926 – Ecuador, Costa Rica (to Chionomus gluciophilus (Muir, 1926), combination by Weglarz and Bartlett 2020: 23.
elphacodes guianensis Muir, 1919 – British Guiana – to Nilaparvata by Kennedy & Bartlett, 2021.
Delphacodes havanae Muir & Giffard, 1924: 37; to Chionomus havanae (Muir & Giffard, 1924) by Fennah 1971a: 323–324.
Delphacodes humilis (Van Duzee, 1907: 48) -See Syndelphax
Delphacodes hyalina Beamer, 1946d: 141; syn with Nothodelphax albocarinatus (Stål, 1858) by Wilson 1992: 85–86
Delphacodes incerta (Van Duzee, 1897: 258); to Javesella incerta (Van Duzee, 1897) by Bartlett et al. 2014: 119.
Delphacodes incurva Beamer, 1948a: 3, 7; to Flavoclypeus incurvus (Beamer 1948a) by Kennedy and Bartlett 2014: 28.
27 Delphacodes indentistyla Dozier, 1926 – USA: MS
Delphacodes indistincta (Crawford, 1914: 619); to Elachodelphax (Elachodelphax) indistinctus (Crawford, 1914); comb. by Hamilton 2002a: 17.
Delphacodes idonea Beamer, 1947: 66 – see Toya
Delphacodes kilmani (Van Duzee, 1897: 253); to Javesella kilmani (Van Duzee, 1897) by Wilson 1992: 97; then to Paraliburnia kilmani (Van Duzee, 1897) by Hamilton 2002a: 22.
28 Delphacodes koebelei Muir & Giffard, 1924 – Mexico, Ecuador
29 Delphacodes kuscheli Fennah, 1955 -Chile, Juan Fernández Islands, Argentina
Delphacodes laminalis (Van Duzee, 1897: 251); to Calligypona laminalis (Van Duzee, 1897) by Kontkanen 1958: 143; then to Muellerianella laminalis (Van Duzee, 1897) by Maw et. al. 2000: 85.
30 Delphacodes lappae Beamer, 1946 – USA: TX
Delphacodes laterana Metcalf, 1943: 458; replacement name for Liburnia lateralisVan Duzee 1897: 253 nec Liburnia striatella var lateralis Fieber 1879; jr. syn. of Muellerianella laminalis (Van Duzee, 1897) by Oman 1947: 212–213.
Delphacodes lateralis (Van Duzee, 1897: 253) (nec Liburnia striatella var. lateralis Fieber 1879); jr. syn. of Muellerianella laminalis (Van Duzee, 1897) by Oman 1947: 212–213.
Delphacodes latidens Beamer, 1948a: 4, 7; to Flavoclypeus latidens (Beamer 1948a) by Kennedy and Bartlett 2014: 29.
31 Delphacodes latus De Remes, Lenikov & Teson, 1979: 322 (asl) – Argentina
Delphacodes lineatipes (Van Duzee, 1897: 255); to Calligypona lineatipes (Van Duzee, 1897) by Kontkanen 1958: 143; then to Nothodelphax lineatipes (Van Duzee, 1897) by Fennah 1965d: 254.
Delphacodes litoralis (Reuter, 1880: 198); to Paraliburnia (Struebingianella) litoralis (Reuter, 1880); comb. by Le Quesne 1964: 57; then to Struebingianella litoralis (Reuter, 1880); status by Nast 1972: 62; then to Paradelphacodes litoralis (Reuter, 1880) by Anufriev 1980: 211.
32 Delphacodes livida Beamer, 1948 – USA: CT
Delphacodes lutea Beamer, 1946d: 142; to Muirodelphax luteus (Beamer, 1946d) by Bartlett et al. 2014: 127.
Delphacodesluteivitta (Walker, 1851: 354); to Spartidelphaxluteivittus (Walker, 1851) by Bartlett and Webb 2014: 32.
Delphacodes lutulenta (Van Duzee, 1897: 252); to Calligypona lutulenta (Van Duzee, 1897) by Kontkanen 1958: 143; then to Kosswigianella (Kosswigianella ) lutulenta (Van Duzee, 1897) by Hamilton & Kwon 2010: 421–422.
Delphacodes lutulentella Muir & Giffard, 1924: 24; to Javesella lutulentella (Muir & Giffard, 1924) by Maw et. al. 2000: 84.
Delphacodes lutulentoides Beamer, 1948a: 5; to Kosswigianella lutulentoides (Beamer, 1948a) by Bartlett et al. 2014: 121.
Delphacodes magna (Crawford, 1914: 627); to Ribautodelphax magna (Crawford, 1914) by Bartlett et al. 2014: 141.
33 Delphacodes marginicornis (Fowler, 1905: 135) – Mexico
34 Delphacodes mcateei Muir and Giffard, 1924 – USA: IL, MD, MO
Delphacodes megadonta Beamer, 1951a: 11; to Aethodelphax megadontus (Beamer, 1951a) by Bartlett & Hamilton 2011: 59–60.
35 Delphacodes mesada Caldwell 1951 – Puerto Rico
Delphacodes montezumae Muir and Giffard, 1924: 27; to Isodelphax montezumae (Muir & Giffard, 1924) by Bartlett et al. 2014: 118.
Delphacodes munda Beamer, 1948c: 116-117; to Kusnezoviellamunda (Beamer, 1948c) by Bartlett et al. 2014: 121,
Delphacodes neocclusa Muir & Giffard, 1924: 22; to Nothodelphax neocclusa (Muir & Giffard, 1924) by Fennah 1965d: 254.
Delphacodes nigra (Crawford, 1914: 624) – see Toya
Delphacodes nigridorsum (Crawford, 1914: 620); to Isodelphax nigridorsum (Crawford, 1914) by Bartlett et al. 2014: 118.
Delphacodes nigrifacies Muir, 1918: 428; to Flavoclypeus nigrifacies (Muir, 1918) by Kennedy and Bartlett 2014: 30.
= Delphacodes xerosa Caldwell, 1951; see Kennedy et al 2012.
Delphacodes nigriscutellata Beamer, 1947: 62; to Caenodelphax nigriscutellata (Beamer, 1947) by Bouchard et al. 2001: 49; then to Flavoclypeus nigriscutellatus by Kennedy and Bartlett 2014: 32.
Delphacodes nigrigaster (Crawford, 1914: 621); syn. with Ribautodelphax albostriatus (Fieber) by Bartlett et al. 2014: 141.
36 Delphacodes nigrinota Beamer, 1951 – USA: TX; Mexico
37 Delphacodes nigripennata Beamer, 1946 – USA: FL, KS, NC
Delphacodes nigripennis (Crawford, 1914: 625) – See Syndelphax
Delphacodes nitens Muir and Giffard, 1924: 27; to Flavoclypeus nitens (Muir and Giffard, 1924) by Kennedy and Bartlett 2014: 33.
Delphacodes obscurella (Boheman, 1847: 53); to Javesella obscurella (Boheman, 1847) by Le Quesne 1964: 57.
Delphacodes occlusa (Van Duzee, 1897: 256); to Nothodelphax occlusa (Van Duzee, 1897) by Fennah 1965d: 254.
Delphacodes opaca Beamer 1948a: 8; to Javesella opaca (Beamer, 1948a) by Maw et. al. 2000: 84.
Delphacodes osborni (Van Duzee, 1897: 250); to Delphacodes campestris by Oman 1947: 210–211 (jr. syn. ofMuirodelphax arvensis (Fitch, 1851) by Hamilton & Kwon 2010: 423.
38 Delphacodes pacifica (Crawford, 1914: 626) – USA: AZ, CA, IL, MO, MS; Mexico (to Chionomus pacificus (Crawford, 1914), combination by Weglarz and Bartlett 2020: 29.)
Delphacodes pallidivitta Fennah, 1945a: 433 – jr. syn ofTagosodes cubanus, see Tagosodes
39 Delphacodes paludicola Muir, 1926 – Ecuador
Delphacodes paransera Beamer, 1948b: 101; to Calligypona paransera (Beamer, 1948b) by Kontkanen 1958: 143; then to Elachodelphax (Aschedelphax) paransera (Beamer, 1948b); comb. by Hamilton & Kwon 2010: 421.
Delphacodes paraparvula Beamer, 1948c: 112 – see Aethodelphax
Delphacodes parvula (Ball, 1902b: 264); to Muirodelphaxparvulus (Ball, 1902b) by Hamilton & Kwon 2010: 423.
40 Delphacodes patruelis (Stal, 1859) – Argentina
Delphacodes pediforma Beamer, 1947: 70; to Yukonodelphax pediforma (Beamer, 1947) by Wilson 1992: 92; then to Elachodelphax (Aschedelphax) pedaforma (Beamer, 1947); comb. and missp. by Hamilton 2002a: 17; also Hamilton & Kwon 2010: 17.
Delphacodes penedetecta Beamer, 1950: 70; to Spartidelphax penedetectus (Beamer, 1950) by Bartlett and Webb 2014: 23.
Delphacodes penelutea Beamer, 1948c: 112; to Muirodelphax peneluteus (Beamer, 1946) by Bartlett et al. 2014: 127.
Delphacodes peneparvula Beamer, 1947: 61-62; to Elachodelphax peneparvula (Beamer, 1947) by Bartlett et al. 2014: 116.
41 Delphacodes penepuella Beamer, 1948 – USA: FL, NY
Delphacodes perusta Beamer, 1947: 62 to Kosswigianella perusta (Beamer, 1947) by Bartlett et al. 2014: 121.
Delphacodes pellucida (Fabricius, 1794: 7); to Calligypona pellucida (Fabricius 1794) by Kontkanen 1958: 143; then to Javesella pellucida (Fabricius, 1794) by Fennah 1963a: 15.
Delphacodes philyra Fennah, 1959: 262; Caenodelphax philyra (Fennah, 1959) by Fennah, 1965: 96; syn. with Caenodelphax teapae by Kennedy and Bartlett 2014: 21.
42 Delphacodes platystylus Muir, 1930 – Argentina
Delphacodes pictifrons Osborn, 1938: 344; syn. with Pissonotus piceus by Bartlett & Deitz 2000: 107.
Delphacodes plenatra Beamer, 1948c: 114-115: to Isodelphax plenatra (Beamer, 1948c) by Bartlett et al. 2014: 118.
Delphacodes propinqua (Fieber, 1866b: 525) (see Metadelphax); to Calligypona propinqua (Fieber, 1866b) by Dlabola 1954: 14, 64 (and by implication China 1954: 165); then to Metadelphax propinqua (Fieber, 1866b) by Wagner 1963: 170; and Toya propinqua (Fieber, 1866b) by Fennah 1964: 142 (from Delphacodes); see also Fennah 1965b: 56 (syn. of Metadelphax by implication) restored to Metadelphax propinqua (Fieber, 1866b) by Ding 2006: 511, 513–514; see also Gonzon & Bartlett, 2008: 242–246.
Delphacodes puella (Van Duzee, 1897: 250) – USA: AL, AR, CA, CT, CO, DC, FL, GA, HI, IA, IL, IN, KS, KY, LA, MA, MD, ME, MO, MS, NC, NE, NH, NJ, NY, OH, OK, OR, PA, TN, VA, WA; Canada: Que., Ont.; Bermuda, Brazil, Cuba, Mexico, Puerto Rico (to Chionomus puellus (Van Duzee, 1897), combination by Weglarz and Bartlett 2020: 31).
43 Delphacodes quadridentis Beamer, 1948 – USA: MS, FL
44 Delphacodes quadrispinosa Muir and Giffard, 1924 – USA: FL; Cuba, Nicaragua (to Chionomus quadrispinosus (Muir & Giffard, 1924), combination by Weglarz and Bartlett 2020: 34).
45 Delphacodes rectangularis (Crawford, 1914: 612) – Nicaragua
46 Delphacodes recurvata Beamer, 1948 – USA: DC, FL, NC
47 Delphacodes reducta (Van Duzee, 1907: 49) – Jamaica
48 Delphacodes rivularis (Germar, 1830) – USA: GA (nomen dubium)
Delphacodes rotundata (Crawford, 1914: 618); syn with Muirodelphax parvulus (Ball, 1902b) by Hamilton & Kwon 2010: 423.
49 Delphacodes saccharicola Muir, 1926 – Ecuador
Delphacodes sagae Beamer, 1946: 141; to Falcotoya sagae (Beamer, 1946d) by Bartlett et al. 2014: 117.
50 Delphacodes sagata (Fowler, 1905: 135) – Mexico, Panama
Delphacodes sagittata Beamer 1947: 64; to Aethodelphax sagittatus (Beamer, 1947) by Bartlett & Hamilton 2011: 61–63.
Delphacodes saileri Beamer 1952: 114; syn. with Javesella simillima (Linnavuori, 1948) by Wilson 1988: 341.
51 Delphacodes saxicola Muir, 1926 – Brazil (Rio de Janeiro)
52 Delphacodes scolochloa Cronin and Wilson 2007 – USA: ND
53 Delphacodes securigera Muir, 1926 – Brazil (Rio de Janeiro)
54 Delphacodes selkirki Fennah, 1955 – Juan Fernández Islands
55 Delphacodes seminigra (Stal, 1854: 246) – Brazil (probably in error USA: CO, FL, Jamaica)
Delphacodes serrata Beamer, 1948a:6-8; to Nothodelphax serrata (Beamer, 1948a) by Bartlett et a;. 2014: 131.
56 Delphacodes shermani (Metcalf, 1923) – USA: NC, KS
57 Delphacodes silvae Beamer, 1946 – USA: TX
58 Delphacodes sitarea De Remes, Lenikov & Teson, 1979 – Argentina
59 Delphacodes spinosus Teson & de Remes Lenicov 1983 – Argentina
60 Delphacodes staminata (Metcalf, 1923) – USA: IL
Delphacodes stejnegeri (Ashmead, 1899: 340); syn. with Javesella pellucida (Fabricius) by Vilbaste 1980: 373.
Delphacodes stramineosa Beamer, 1948a: 5 – see Yukonodelphax
61 Delphacodes stricklandi Metcalf, 1946 – CAN: AB, MB, SK
62 Delphacodes sucinea Beamer, 1948 – USA: FL, MS
elphacodes tenae Muir, 1926 – Ecuador (to Chionomus tenae (Muir, 1926), combination by Weglarz and Bartlett 2020: 36.
Delphacodes teapae (Fowler, 1905: 135); to Caenodelphax teapae (Fowler, 1905); comb. by Fennah 1965a: 96.
63 Delphacodes trimaculata Beamer, 1948 – USA: CT, DC, IL, MD, VA
64 Delphacodes truncata Beamer, 1948 – USA: FL, GA, LA, MO, MS, VA
65 Delphacodes turgida Beamer, 1948 – USA: FL, GA
Delphacodes uhleri Muir and Giffard, 1924 – jr. syn of Syndelphax alexanderi See Syndelphax
Delphacodes unda (Metcalf, 1923: 207) (See Muirodelphax); syn. with Delphacodes campestris (Van Duzee) by Metcalf 1949: 56; removed from syn. as Muirodelphax unda (Metcalf) by Hamilton & Kwon 2010: 423.
Delphacodes uniformis Metcalf 1943: 521 (incertae sedis) (n. n. for D. unicolor Walker, 1851: 351, nec Herrich-Schäffer 1835) (type specimen is nymph and not delphacid, see Muir 1919)
66 Delphacodes univittata (Berg, 1879: 189) – Argentina
Delphacodes vaccina Caldwell, 1951 – Puerto Rico (synonymy with Chionomus dissipatus (Muir, 1926) by Weglarz and Bartlett 2020: 19.
Delphacodes venusta Beamer, 1948c: 115; to Nothodelphax venusta (Beamer, 1948c) by Hamilton 2002a: 22.
67 Delphacodes waldeni (Metcalf, 1923) – USA: CT
Delphacodes wetmorei Muir & Giffard, 1924: 30; to Metadelphax by Gonzon & Bartlett 208: 247.
68 Delphacodes xerophila Muir, 1926 – Ecuador
Delphacodes xerosa Caldwell, 1951 (in Caldwell & Martorell 1951); synonymy with Delphacodes nigrifaces by Kennedy et al., 2012: 405.
Plant associations
Delphacodes s.s.
Delphacodes capnodes (Scott) – Carex fuscula var. pseudextensa (Kük.) Kük. (As Carex riparia Curtis) (River sedge, Cyperaceae)
Delphacodes schinias Asche et al. – Cyperus sp. (Cyperaceae), Phragmites sp. (Poaceae)
Delphacodes venosus on British Bugs it is noted that this species is associated with “Sphagnum, sedges or rushes, where it is commonly found near their base. The host itself is a range of grasses including Deschampsia cespitosa, Festuca and Mollinia [sic] caerulea”
Delphacodes s.l.
Delphacodes arcuata Beamer – Manihot esculenta Crantz (cassava; as Manihot utilissima Pohl; Euphorbiaceae)
Delphacodes bellicosa Muir and Giffard – Paspalum distichum L. (knotgrass, Poaceae)
Delphacodes cerberus Fennah –Drimys confertifolia Phil. (Winteraceae)
Delphacodes muirella Metcalf – Corokia sp. (Argophyllaceae), Metrosideros Banks ex Gaertn (Myrtaceae), Blechnum L. (Blechnaceae)
Delphacodes nigerrima Ishihara – Murdannia keisak (Hassk.) Hand.-Maz. (Commelinaceae)
Delphacodes puella (Van Duzee) – Galinsoga parviflora Cav. (Asteraceae), Panicum capillare L. (witchgrass, Poaceae)
Delphacodes stricklandi Metcalf – Amaranthus sp. (Amaranthaceae) (given as A. “elapious”, but I can find no such species)
[Hosts from Wilson et al. 1994; names check on USDA Plants OR Tropicos].
Economic Importance
While most species of “Delphacodes” are obscure or rarely encountered, a variety of species have been found in agricultural crops and may serve as potential vectors of plant disease.Delphacodes kuscheli is a vector of Mal de Rio Cuarto (MRCV) in corn in South America.
Delphacodes catilina Fennah, 1958 (as “Toya catilina“) is a vector of Maize mosaic nucleorhabdovirus (VIDE Rhabdoviridae) (see also here) (as “African Cereal Streak Virus”) (Harder and Bakker 1973; Abo and Sy 1997).
Recognition
Descriptive details still to come – Below are illustrations from Asche & Remane 1983 on Delphacodes in the strict sense.
Interpretations of Delphacodes mulsanti (from Asche and Remane 1983)
Delphacodes fascia (paratype)
Delphacodes fascia (from Asche and Remane 1983)
Delphacodes fascia paratype from Asche and Remane 1983
Delphacodes nastaci (from Asche and Remane 1983)
Delphacodes nastaci (from Asche and Remane 1983)
Delphacodes schiniae and D. framarib (from Asche and Remane 1983)
Delphacodes schiniae and D. framarib (from Asche and Remane 1983)
Delphacodes schiniae and D. framarib (from Asche and Remane 1983)
Delphacodes schiniae and D. framarib (from Asche and Remane 1983)
Photos by Kimberley Shropshire (University of Delaware, Department of Entomology and Wildlife Ecology) unless otherwise indicated.
Some fairly awful pictures of Delphacodes mulsanti (Det. by Wagner) – posted because this is the type species.
Delphacodes mulsanti (det. Wagner, a poor specimen, but posted because this is the type species)
Delphacodes audrasi (also Delphacodes s.s.).
Delphacodes capnodes (Det. Wagner)
Delphacodes capnodes (also Delphacodes s.s.). (Det Wagner – also an awful specimen)
Delphacodes capnodes FLOW from Gernot Kunz.
Delphacodes capnodes FLOW from Gernot Kunz.
New World speciesof uncertain status
Delphacodes acuministyla
Delphacodes acuministyla
Delphacodes agropyri
Delphacodes angulata
Delphacodes apicata
Delphacodes aterrima
Delphacodes aterrima (holotype)
Delphacodes balli
Delphacodes bocana
Delphacodes caerulata (female, the male paratype I had photographed turned out to be misidentified)
Delphacodes darwini
Figs. 25-32 (Campodonico 2017). Delphacodes darwini, brachypterous male. 25.- Habitus, dorsal view. 26.- Habitus, lateral view. 27.- Head, frontal view. 28.- Pygofer, caudal view. 29.- Pygofer, lateral view. 30.- Anal tube, lateral view. 31.- Right paramere, caudal view. 32.- Aedeagus, lateral view. Figs. 33-41.- Delphacodes kuscheli, brachypterous male. 33.- Habitus, dorsal view. 34.- Habitus, lateral view. 35.- Head, frontal view. 36.- Pygofer, caudal view. 37.- Pygofer, lateral view. 38.- Anal tube, lateral view. 39.- Anal tube, caudal view. 40.- Right paramere, caudo-lateral view. 41.- Aedeagus, lateral view.
Figs. 33-41 (Campodonico 2017). Delphacodes kuscheli, brachypterous male. 33.- Habitus, dorsal view. 34.- Habitus, lateral view. 35.- Head, frontal view. 36.- Pygofer, caudal view. 37.- Pygofer, lateral view. 38.- Anal tube, lateral view. 39.- Anal tube, caudal view. 40.- Right paramere, caudo-lateral view. 41.- Aedeagus, lateral view.
Delphacodes lappae (macropter)
Delphacodes lappae (brachypter)
Delphacodes livida
Delphacodes mcateei (I recall this species being mostly brachypterous but still need good images of brachypters)
Delphacodes mesada
Delphacodes nigrinota
Delphacodes nigripennata
Delphacodes recurvata
Delphacodes sucinea
Delphacodes sucinea
Delphacodes trimaculata paratype
Delphacodes trimaculata (male)
[for some reason this page was truncated and I lost all beyond this point; will try to fix, but I think I will have to recreate from scratch]
Online resources
[still to come]
Molecular resources
[still to come]
Selected Literature
[these need to be aligned with the above]
Anufriev, G. A. 1977. Delphacids (Homoptera Auchenorrhyncha) of the Kurile Islands Fauna. Zoologicheskii Zhurnal 56(6): 855-869. [In Russian.]
Asche, M. 1982. Maculidelphax gen. nov., eine neue Gattung fur Calligypona maculipennis Linnavuori, 1962 (T.g.) und Delphacodes epimelas Fennah, 1958 (Homoptera Cicadina Delphacidae). Marburger Entomologische Publikationen 1(6): 191-210.
Asche, M. 1985. Zur Phylogenie der Delphacidae Leach, 1815 (Homoptera: Cicadina: Fulgoromorpha). Marburger Entomologische Publikationen 2(1), volume 1 pp. 1-398, volume 2 pp. 399-910.
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