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Contents
Family Delphacidae Leach, 1815
Subfamily Delphacinae Leach, 1815
Tribe Delphacini Leach, 1815
Genus Javesella Fennah, 1963: 15.
Type species (in original combination): Fulgora pellucida Fabricius, 1794: 7.
Synonyms:
= Weidnerianella Wagner, 1963 (Type species Fulgora pellucida Fabricius, 1794; syn. by Wagner 1966: 91; see also Le Quesne 1964: 57.
= Javasella Fennah, 1963a; missp. by Maw et al. 2000: 84 and Hamilton 2002a: 17.
Subgenera:
All species have not been assigned to subgenera. The two described subgenera are:.
Subgenus Haffnerianella Wagner 1966 (type species Delphacodes stali Metcalf 1943, replacement name for the unavailable Delphax bohemani Stål, 1858, nec. Stål, 1854)
Subgenus Javesella Fennah, 1963.
Distribution
Widespread in Holarctic, especially north; some species circumboreal.
Recognized species
There are 23 species of Javesella as follows (I still need to add synonymies for Paleartic species):
Subgenus Javesella
1 Javesella alpina (Sahlberg, 1871) – Finland, Sweden, Norway, Kanin Peninsula (Arctic regions of Fennoscandia and Siberia), North European Russia
= Liburnia alpina
Javesella alpina tota (Sahlberg, 1871) (the status of this name is not clear to me)
= Liburnia alpina tota Sahlberg, 1871
Delphacodes alpina tota (Sahlberg, 1871)
Note this subspecies and species listed as a junior synonym of Javesella forcipata by Nast 1972: 67. Ossiannilsson (1978: 200) treated alpina as a good species (but did not mention the subspecies, that I see).
2 Javesella azorica Remane, 1979: 2 – Azores
3 Javesella bottnica Hulden, 1974 – Finland, Russia
4 Javesella discolor (Boheman, 1847) – USA: Alaska; Canada: Yukon, Northwest territories; widespread in Europe, Russia, Mongolia, Northern Africa; East Palaearctic; Austria, Belgium, Bulgaria, Czech Republic, Denmark, Estonia, Finland, France, Germany, Great Britain, Greece, Ireland, Italy, Latvia, Lithuania, Netherlands, North European Russia, Norway, Poland, Romania, Sicily, Slovakia, Sweden, Switzerland, Yugoslavia
= Delphax discolor Boheman, 1847: 61.
= Liburnia discolor (Boheman, 1847); comb. by Scott, 1870: 75.
= Delphacodes discolor (Boheman, 1847); comb. by Le Quesne, 1964: 57.
= Javesella discolor (Boheman, 1847); comb. by Le Quesne, 1964: 57.
= Delphax fuscipennis Kirschbaum 1868; syn. By Wagner 1939.
= Delphax nitidula Kirschbaum 1868; syn. By Wagner 1939.
= Delphax patens Kirschbaum 1868; syn. By Wagner 1939.
= Delphax similis Kirschbaum 1868; syn. By Wagner 1939.
5 Javesella dubia (Kirschbaum, 1868) – Widespread in Palearctic; reported, e.g., Azores, Morocco, Altai, Kazakhstan, Uzbekistan; East Palaearctic; Albania, Austria, Belgium, Bosnia and Herzegovina, Bulgaria, Central European Russia, Corsica, Croatia, Cyprus, Czech Republic, Denmark, Estonia, Finland, France, Germany, Great Britain, Greece, Hungary, Ireland, Italy, Japan (Hokkaido), Kriti (Crete), Latvia, Lebanon, Lithuania, Macedonia, Madeira Is., Moldova, Netherlands, Norway, Poland, Romania, Russia (Altay, Kamchatka, Kuril Islands, Primorye, Sakhalin, Siberia (Krasnoyarsk)), Sardinia, Slovakia, Spain, Sweden, Switzerland, Turkey, Yugoslavia
= Delphax dubia Kirschbaum, 1868
= Delphax herrichi Kirschbaum, 1868; synonym by Wagner 1939: 125.
= Delphax nitidipennis Kirschbaum, 1868; synonym by Wagner 1939: 125.
= Liburnia difficilis Edwards, 1888; synonym by Wagner 1939: 125; Le Quesne (1960: 43).
= Liburnia pargasensis Reuter, 1880; synonym by Wagner 1963.
6 Javesella forcipata (Boheman, 1847) – Widespread in Palearctic, except south; East Palaearctic; Austria, Belgium, Bosnia-Herzegovina, Bulgaria, Czech Republic, Denmark, Estonia, Finland, France, Germany, Great Britain, Greece, Hungary, Ireland, Italy, Japan (Hokkaido, Honshu), Kazakhstan, Latvia, Lithuania, Mongolia, Netherlands, Norway, Poland, Portugal, Romania, Russia (Buryatiya, Chita, Khabarovsk, Primorye, Tuva, Siberia (Krasnoyarsk); North European Russia, Central European Russia), Slovakia, Sweden, Switzerland, Turkey, Ukraine, Yugoslavia
= Delphax forcipata Boheman, 1847, original combination.
= Delphacodes angustata Metcalf, 1943; syn. by Wagner 1963: 176.
= Delphacodes dorsigera Metcalf, 1943; syn. by Wagner 1963: 176.
= Delphacodes rufescens Metcalf, 1943; syn. by Wagner 1963: 176.
7 Javesella obscurella (Boheman, 1847) – USA: Alaska, Canada: Yukon, Northwest Territories; widespread Palearctic (Eurosiberian), Mongolia; East Palaearctic; Austria, Belgium, Bosnia and Herzegovina, Bulgaria, Czech Republic, Denmark, Estonia, Finland, France, Germany, Great Britain, Greece, Hungary, Ireland, Italy, Latvia, Lithuania, Moldova, Netherlands, Norway, Poland, Portugal, Romania, Russia (North European Russia, Central European Russia, South European Russia), Sardinia, Slovakia, Slovenia, Sweden, Yugoslavia
= Delphax obscurella Boheman, 1847: 53
= Liburnia obscurella Boheman, 1847; comb. by Scott, 1870: 28.
= Delphacodes obscurella (Boheman, 1847); comb. by China, 1938: 197.
= Javesella obscurella (Boheman, 1847); comb. by Le Quesne, 1964: 57.
= Delphacodes fuscotestacea Metcalf, 1943
= Liburnia discreta Edwards, 1888
8 Javesella pellucida (Fabricius, 1794) [Metcalf 1943: 482] – Holarctic; widespread and common, especially north. In New World, reported from Alaska, throughout Canada, widespread in the US, especially northern states, but reported from Cuba and Puerto Rico. In Palearctic, northern Europe to Northern Africa Including the Azores) east to Mongolia and Japan. Also reported Iceland. USA: AK, AZ, CA, CO, CT, DE, ID, IL, MA, MD, ME, MI, MO, NC, ND, NH, NJ, NM, NV, NY, OH, OR, PA, TN, UT, VT, WA, WI, WY; CAN: AB, BC, LB, MB, NB, NF, NS, NT, NU, ON, QC, SK, YK; Cuba, Puerto Rico; Palearctic: Algeria, Austria, Belgium, Bosnia and Herzegovina, Bulgaria, Croatia, Czechoslovakia, Denmark, Estonia, Finland, France, Germany (inc. Baltrum Island), Great Britain, Greece, Hungary, Iceland, Ireland, Iran, Italy (inc. Sardegna), Japan (Hokkaido, Honshu), Kazakhstan, Kyrgyzstan, Lebanon, Libya, Lithuania, Macedonia, Mongolia, Morocco, Netherlands (inc. Nijmegen as “Batavia”), Norway, Poland, Portugal, Romania, Russia (Altay, Buryatiya, Chita, Irkutsk, Kamchatka, Khabarovsk, Kuril Islands, Magadan, Primorye, Sakhalin, Tuva; Siberia (Krasnoyarsk); Caucasus, Kamchatka, Sakhalin Island, Siberia, Transcaucasia), Sardinia, Scandinavia, Scotland, Slovakia, Spain (inc. Canary Islands), Sweden, Switzerland, Turkey, Turkestan, Ukraine, Uzbekistan, Wales, Yugoslavia.
= Fulgora pellucida Fabricius, 1794: 7.
= Fulgora striata Fabricius,1794; syn. with Delphax dispar Fallén, 1806 by Zetterstedt, 1828: 548; syn. with Delphax pellucida by Flor, 1861: 65.
= Fulgora marginata Fabricius, 1794; syn. with Delphax dispar Fallén, 1806 by Fallén, 1806: 126; syn. with Delphax pellucida by Herrich-Schäffer, 1836: 18.
= Fulgora flavescens Fabricius, 1794; syn. with Delphax dispar Fallén, 1806 by Fallén, 1806: 126; syn. with Delphax pellucida by Flor, 1861: 65.
= Cicada pellucida (Fabricius, 1794); comb. by Weber, 1795: 146.
= Delphax pellucida (Fabricius, 1794); comb. by Fabricius, 1803 :84.
= Delphax striata (Fabricius, 1794); comb. by Fabricius, 1803: 84.
= Delphax marginata (Fabricius, 1794); comb. by Fabricius, 1803: 84.
= Delphax flauescens [sic] (Fabricius, 1794); comb. by Fabricius, 1803: 84.
= Asiraca pellucida (Fabricius, 1794); comb. by Latreille, 1804: 318.
= Asiraca striata (Fabricius, 1794); comb. by Latreille, 1804: 318.
= Asiraca marginata (Fabricius, 1794); comb. by Latreille, 1804: 318.
= Asiraca flavescens (Fabricius, 1794); comb. by Latreille, 1804: 318.
= Delphax dispar Fallén, 1806; syn. with Delphax pellucida by Flor, 1861: 65..
= Delphax suturalis Curtis, 1837; syn. with Delphax pellucida by Flor, 1861: 65.
= Delphax dubia Kirschbaum, 1868 (nec. Panzer, 1796); syn. with Liburnia pellucida (Fabricius, 1794) by Fieber, 1872: 5.
= Delphax piceola Kirschbaum, 1868; syn. with Delphax pellucida (Fabricius, 1794) by Puton, 1886: 73.
= Liburnia pellucida (Fabricius, 1794); comb. by Stål, 1869: 117.
= Liburnia striata (Fabricius, 1794); comb. by Stål, 1869: 117.
= Liburnia marginata (Fabricius, 1794); comb. by Stål, 1869: 117.
= Liburnia flavescens (Fabricius, 1794); comb. by Stål, 1869: 117.
= Achorotile pellucida (Fabricius, 1794); comb. by Oshanin, 1870: 48.
= Liburnia suturalis (Curtis, 1837); comb. by Fieber, 1872: 5.
= Delphax pellucida dispar (Fallén, 1806); comb. by Mayr, 1884 :20.
= Liburnia pellucida dispar (Fallén, 1806); comb. by Melichar, 1896: 71.
= Liburnia yezoana Matsumura, 1900; syn. by Muir and Giffard, 1924: 20.
= Embolophpora [sic.] pellucida (Fabricius, 1794); comb. by Kirkaldy, 1901: 338.
= Delphax yezoana (Matsumura, 1900); comb. by Oshanin, 1907: 330.
= Megamelus pellucidus (Fabricius, 1794); comb. by Crawford, 1914: 607, 615.
= Liburnia arvensis (Fitch, 1856); syn. by Crawford 1914: 615 (error).
= Liburnia semicinctus Van Duzee, 1917; syn. with Delphacodes pellucida by Muir and Giffard, 1924: 20.
= Delphacodes pellucida (Fabricius, 1794); comb. by Muir and Giffard, 1924: 20.
= Javesella pellucida (Fabricius, 1794); comb. by Fennah, 1963: 15.
= Delphacodes stejnegeri (Ashmead, 1899); syn. by Vilbaste, 1980: 373.
9 Javesella salina (Haupt, 1924) – Eastern Palearctic (Central, Eastern, and Southeastern Europe, Asia Minor), Mongolia; East Palaearctic; Bulgaria, Czech Republic, Estonia, Germany, Hungary, Lithuania, Poland, Slovakia, Sweden
= Delphax salina Haupt, 1924
= Liburnia juncea Haupt, 1935; new name for Delphax discolor Fieber 1866 [nec. Boheman]; syn with Javesella salina (Haupt) by Wagner 1963: 176.
= Delphacodes juncea (Haupt); comb. by Metcalf 1943: 456.
= Javesella salina (Haupt, 1924); comb. by Wagner 1963: 176.
10 Javesella simillima (Linnavuori, 1948) – USA: Alaska; Canada: Manitoba, Northwest Territories, Yukon; East Palearctic – Arctic regions; Finland, Estonia, Finland, Germany, Poland, Romania, Russia (North European Russia,), Germany, East Fennoscandia.
= Calligypona simillima Linnavuori, 1948.
= Javesella simillima (Linnavuori, 1948); comb. by Nast, 1972: 68.
= Delphacodes saileri Beamer 1952; syn. by Wilson, 1988: 341.
Subgenus Haffnerianella
11 Javesella stali (Metcalf, 1943) – Northern, eastern and central Europe, Eastern Siberia; Austria, Czech Republic, Estonia, Finland, France, Germany, Hungary, Italy, Latvia, Lithuania, Mongolia, North European Russia, Poland. Slovakia, Sweden.
= Delphacodes stali Metcalf, 1943
Not placed to subgenus
12 Javesella arcanastyla (Beamer, 1948) – USA: Alaska, Michigan, Washington, Wisconsin, Wyoming; Canada: British Columbia, Saskatchewan, Yukon.
= Delphacodes arcanastyla Beamer, 1948: 101.
= Javesella arcanastyla (Beamer, 1948); comb. by Wilson, 1988: 341.
13 Javesella atrata (Osborn, 1938) – USA: Alaska, Connecticut, Illinois, New Hampshire, New York, Ohio; Canada: Alberta Manitoba, Ontario, Quebec, Yukon.
= Delphacodes atrata Osborn, 1938: 344.
= Javesella atrata (Osborn, 1938); comb. by Wilson, 1988: 341.
14 Javesella badia Anufriev, 1988: 422 (In Anufriev and Emeljanov 1988) – Russia (Prim., Sikhote Alin Reserve, Upper Nantsa)
15 Javesella beringiaca Emeljanov, 1992 – Canada: Manitoba, Saskatchewan, Yukon; Russia (Kamchatka, Magadan, Sakhalin Regions)
16 Javesella compacta Mitjaev, 1987 – Russia (Kazakhstan)
17 Javesella dolera (Spooner, 1912) – USA: Illinois, New York, Virginia
= Liburnia dolera Spooner, 1912: 240.
= Megamelus dolerus (Spooner, 1912); comb. by Crawford, 1914: 630.
= Delphacodes dolera (Spooner, 1912); comb. by Metcalf, 1928: 182.
= [Javesella] dolera (Spooner, 1912); comb. by Maw et. al., 2000: 84.
18 Javesella ila Wilson, 1992 – Canada: Yukon; USA: New Hampshire
19 Javesella incerta (Van Duzee, 1897) – Canada: Quebec; USA: Michigan, New York, Ohio, Pennsylvania
= Liburnia incerta Van Duzee, 1897: 227, 258.
= Dicranotropis incerta (Van Duzee, 1897); comb. by Muir & Giffard 1924: 17.
= Delphacodes incerta (Van Duzee, 1897); comb. by Metcalf, 1928: 182; Oman, 1947: 210-211.
= Javesella incerta (Van Duzee, 1897); comb. by Bartlett et al. 2014: 119.
20 Javesella lutulentella (Muir & Giffard, 1924) – USA: California; Canada: British Columbia.
= Delphacodes lutulentella Muir and Giffard, 1924: 24.
= Javesella lutulentella Muir and Giffard, 1924 comb. by Maw et. al., 2000: 84.
21 Javesella nuchtica Dlabola, 1967 – Mongolia
22 Javesella opaca (Beamer, 1948) – CAN: Ontario, Quebec; USA: Alabama, Connecticut, Georgia, Massachusets, North Carolina, New York, Pennsylvania, South Carolina, Tennessee, Virginia
= Delphacodes opaca Beamer 1948: 8, 10.
= Javesella opaca (Beamer, 1948); comb. by Maw et. al., 2000.
23 Javesella selengica Dlabola, 1970 – Mongolia
Plant associations
(incomplete)
Javesella alpina – Geranium sp., Vaccinium myrtillus, Trollius sp., Dryopteris sp., Phegopteris sp.
Javesella discolor – on grasses: Deschampsia cespitosa, D. flexuosa, Calamagrostis canescens, and others; probably also on sedges (Carex, Scirpus) and rushes (Juncus, Luzula).
Javesella dubia: on grasses, mainly Agrostis stolonifera, and A. capillaris.
Javesella forcipata – on grasses: Deschampsia cespitosa (L.) P. Beauv., Holcus mollis L., probably also Poa trivialis L., Poa annua L. , Calamagrostis canescens (Weber) Roth, Agrostis stolonifera L.
Javesella obscurella – on grasses, mainly Alopecurus geniculatus L., Glyceria fluitans (L.) R. Br. .
Javesella opaca – Polytrichum commune Hedw. (Polytrichaceae) (Wheeler 2003, common hair-cap moss).
Javesella pellucida – Polyphagous, mostly grasses (Festuca, Elymus, Dactylis, Poa, Deschampsia, Agrostis, Lolium, Calamagrostis, Phleum, and others); less commonly on sedges, rushes, and horsetails. Schedonorus arundinaceus (Schreb.) Dumort., nom. cons. (
Javesella salina: reported on the rush Juncus gerardii, and the grasses Puccinellia distans and Briza media.
Javesella simillima – Eriophorum angustifolium; also reported from Carex rostrata.
Javesella stali: Equisetum arvense, E. palustre (horsetail)
Sources for host information: See Holzinger et al. (2003); see also Wilson et al. (1994) and FLOW.
Predators /parasites
Parasitism: ectoparasite
Abrolophus norvegicus (Thor, 1900) (Acari, Trombidiformes, Erythraeidae); see Sömermaa (1973: 196), Stroinski et al. (2013: 196) on Javesella pellucida.
Parasitism: parasitoid
Charletonia cardinalis (c. l. koch, 1837) (C. L. Koch, 1837) (Acari, Trombidiformes, Erythraeidae) ; see Southcott (1966), Stroinski et al. (2013: 197) on Javesella pellucida.
Gonatopus lindbergi Heikinheimo (Insecta, Hymenoptera, Dryinidae) on Javesella pellucida.
Elenchus tenuicornis (Kirby, 1815) (Elenchidae: Strepsiptera) on Javesella pellucida (Sheppard 2014).
Halictophagus silwoodensis Waloff, 1981 (Strepsiptera: Halictophagidae) in Javesella discolor (Bland 1998)
Panstenon oxylus (Walker) (Pteromalidae) on Javesella pellucida (Graham 1969: 94); Mesopolobus graminum (Hardh 1950) (Graham 1969: 663).
Economic Importance
Several species reported as vectors of plant disease in small grains.
Javesella discolor (Oats) vectors Oat sterile dwarf virus (OSDV)
Javesella dubia (tall oat-grass) vectors Oat sterile dwarf virus (OSDV) and European wheat striate mosaic virus (EWSMV)
Javesella obscurella (oats, wheat) vectors Oat sterile dwarf virus (OSDV) and European wheat striate mosaic virus (EWSMV)
Javesella pellucida (barley, maize, oats, tall oat-grass, wheat) vectors Oat sterile dwarf virus (OSDV) and European wheat striate mosaic virus (EWSMV), and Maize rough dwarf virus (MRDV) (Serjeant 1967, Harpaz 1972)
Recognition
A common and widespread genus. Species recognition is sometimes difficult but usually can be managed based on male genitalia. Species recognition for females is difficult and may not always be possible with confidence. Javesella pellucida is the most commonly encountered Javesella in North America. This species can be quite variable in superficial external appearance.
Members of the genus are usually dark colored; the parameres of the male genitalia are widely diverging, narrow and sinuate; processes on segment X are closely approximated (vs. separated in Nothodelphax).
Keys to species for Palearctic taxa can be found in Holzinger et al. (2003) and Anufriev and Emeljanov (1988)
Javesella atrata (Photos by Kimberley Shropshire, Department of Entomology, University of Delaware)
Javesella beringiaca (probably)
Javesella discolor
Javesella dolera
Javesella dolera
Javesella dubia
Javesella lla
Javesella obscurella
Javesella obscurella
Javesella pellucida
Javesella simillima
Molecular Resources
At this time, Genbank has data for Javesella obscurella and Javesella pellucida here. Bold has barcode sequence data for Javesella pellucida here. Urban et al. (2010) sequence 4 genetic loci (18S rDNA, 28S rDNA, wingless, and cytochrome oxidase I) for Javesella pellucida and J. dubia.
Selected References
(May be incomplete, still working on synonymy, plant association and distribution record literature)
Ammar, E. D. 1975. Effect of European wheat striate mosaic, acquired by feeding on diseased plants, on the biology of its planthopper vector Javesella pellucida. Annals of Applied Biology 79: 195-202.
Anufriev, G. A. and A. M. Averkin. 1982. On delphacids of the Komsomolsky Reserve (Khabarovsk Region, USSR) (Insecta, Homoptera, Auchenorrhyncha, Delphacidae). Faunistische Abhandlungen, Staatliches Museum fur Tierkunde in Dresden (1981) 9(6): 95-100.
Anufriev, G. A. and A. F. Emeljanov. 1988. Volume II: Homoptera and Heteroptera. In: P.A. Lehr (ed.). Keys to the Insects of the Far East of the USSR in Six Volumes, Transliteration of the Russian title: Opredelitel’ nasekomykh Dal’nego Vostoka SSSR v shesti tomakh. Vol. 2. Ravnokrylye i poluzhestkokrylye. In: P.A. Lehr (ed.). Keys to the Insects of the Far East of the USSR in Six Volume. Nauka, Leningrad.
Asche, M. 1982. Beitrage zur Delphaciden-Fauna Ungarns (Homoptera Cicadina Delphacidae). Marburger Entomologische Publikationen 1(7): 139-154.
Asche, M. 1982. Beitrage zur Delphaciden-Fauna der Turkei (Anatolien) (Homoptera Cicadina Delphacidae). Marburger Entomologische Publikationen 1(7): 71-98.
Asche, M. 1982. Beitrage zur Delphaciden-Fauna Jugoslawiens und Bulgariens (Homoptera Cicadina Delphacidae). Marburger Entomologische Publikationen 1(7): 99-138.
Asche, M. and H. Hoch. 1982. Beitrage zur Delphaciden-Fauna Griechenlands II (Homoptera Cicadina Delphacidae). Marburger Entomologische Publikationen 1(7): 37-70.
Asche, M. and H. Hoch. 1986. Cixiidae and Delphacidae (Homoptera – Fulgoroidea) from Lebanon – a preliminary synopsis. Pp. : 18-19. In: Proceedings of the 2nd International Congress Concerning the Rhynchota Fauna of Balkan and Adjacent Regions. 18-22 Aug. 1986. Mikrolimni – Prespa, Greece.
Asche, M. and R. Remane. 1982. Beitrage zur Delphaciden-Fauna Griechenlands I (Homoptera Cicadina Delphacidae) Marburger Entomologische Publikationen 1(6): 231-290.
Ashmead, W. H. 1899. Homoptera. In: Reports upon the insects, spiders, mites and myriapodes collected by Dr. L. Stejneger and mr. G.E.H. Barrett-Hamilton on the Commander Islands. The fur seals and fur-seal islands of North Pacific Ocean 4: 328-351. [Delphax stejnegeri Ashmead, 1899].
Barringer, L. E. and C. R. Bartlett. 2018. Pennsylvania planthoppers (Hemiptera: Auchenorrhyncha: Fulgoroidea): relative abundance and incidental catch using novel trapping methods. Insecta Mundi 0661: 1–31.
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Beamer, R. H. 1948. Some new species of Delphacodes (continued) part III. Journal of the Kansas Entomological Society 21(1): 1-10.
Beamer, R. H. 1952. One old and five new species of delphacine fulgorids (Homoptera: Fulgoridae). Journal of the Kansas Entomological Society 25(3): 111-115.
Bland, K. P. 1998. Halictophagus silwoodensis Waloff, 1981 (Strepsiptera: Halictophagidae) confirmed as a Scottish Insect. The entomologist’s record and journal of variation 110(1-12): 296-297. [Javesella discolor, etc noted]
Boheman, C. H. 1847. Nya Svenska Homoptera. Ofversigt af Kongliga Svenska Vetenskaps-Akademiens Forhandlingar 1847: 263-266. [Delphax palliata Boheman, 1847]
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De Vrijer, P.W.F. 1984. Variability in calling signals of the planthopper Javesella pellucida (F. ) (Homoptera: Delphacidae) in relation to temperature, and consequences for species recognition during distant communication. Netherlands Journal of Zoology 34(3): 388-406.
De Vrijer, P.W.F. 1986. Species distinctiveness and variability of acoustic calling signals in the planthopper genus Javesella (Homoptera: Delphacidae). Netherlands Journal of Zoology 36(1): 162-175.
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